How does vigilance fit into animal's time budgets?
What factors regulate which animal(s) are vigilant at any given moment?
How do proximity of potential predators, experience of the animals in their habitat, and the "culture" of flocks or herds affect time investment in vigilance?
Does vigilance form part of a selfish-herd strategy, or does it better fit reciprocal altruism or kin-selection models?
Body size and predator saturation
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